- The 10% rule
- Only about 10% of the energy at one trophic level is passed to the next; the rest (~90%) is lost as heat through cellular respiration.
- Trophic level
- A feeding position in a food chain: producers, primary consumers, secondary consumers, etc. Energy decreases at each higher level.
- Producer (autotroph)
- An organism that makes its own food, usually by photosynthesis (plants, algae); the base of the food chain.
- Primary consumer
- A herbivore that eats producers (the second trophic level).
- Secondary consumer
- A carnivore that eats primary consumers (the third trophic level).
- Decomposer
- An organism (bacteria, fungi) that breaks down dead matter and recycles nutrients back into the ecosystem.
- Food web
- A network of interconnected food chains showing all the feeding relationships in a community.
- Gross primary productivity (GPP)
- The total rate at which producers capture and store energy through photosynthesis.
- Net primary productivity (NPP)
- GPP minus the energy producers use for their own respiration; the energy available to consumers.
- Keystone species
- A species with a disproportionately large effect on its community; removing it dramatically changes the ecosystem (e.g., sea otters).
- Biogeochemical cycle
- The movement of an element through the living and nonliving parts of an ecosystem (carbon, nitrogen, phosphorus, water).
- Carbon cycle
- Photosynthesis removes CO₂ from the air; respiration, decomposition, and combustion return it. Oceans, soils, and fossil fuels are reservoirs.
- Nitrogen fixation
- Conversion of inert N₂ gas into ammonia (NH₃/NH₄⁺) usable by organisms — done by bacteria, lightning, and the Haber-Bosch process.
- Nitrification
- The bacterial conversion of ammonium (NH₄⁺) to nitrite (NO₂⁻) and then nitrate (NO₃⁻), the form plants use most easily.
- Denitrification
- The bacterial conversion of nitrate (NO₃⁻) back into N₂ gas, returning nitrogen to the atmosphere.
- Phosphorus cycle
- Moves phosphorus through rock, soil, and water with NO atmospheric (gas) phase; phosphorus often limits aquatic plant growth.
- Limiting nutrient
- The nutrient in shortest supply that caps growth; nitrogen often limits land plants, phosphorus often limits aquatic ones.
- The water cycle
- Water moves through evaporation, transpiration, condensation, precipitation, runoff, and infiltration into aquifers.
- Photosynthesis
- Process where producers use sunlight, CO₂, and water to make glucose and release oxygen; it stores energy and removes CO₂.
- Cellular respiration
- Process where organisms break down glucose with oxygen to release energy, returning CO₂ to the atmosphere.
- Energy pyramid
- A diagram showing energy decreasing at each higher trophic level, with producers at the wide base.
- Biome
- A large region defined by its climate and characteristic plant and animal communities (e.g., tundra, desert, rainforest).
- Biodiversity
- The variety of life at three levels — genetic, species, and ecosystem diversity. Higher diversity = more resilience.
- Species richness
- The number of different species in a community.
- Species evenness
- How balanced the populations of different species are in a community.
- Genetic diversity
- The variety of genes within a species; greater diversity helps a species adapt to change and resist disease.
- Ecosystem services
- Free benefits ecosystems provide: pollination, water filtration, flood control, climate regulation, and food.
- Ecological succession
- The gradual, predictable change in a community over time after a disturbance, ending in a climax community.
- Primary succession
- Succession that begins on bare rock with no soil (after a glacier or lava flow); pioneer species are lichens and mosses.
- Secondary succession
- Succession that rebuilds on existing soil after a disturbance (fire, abandoned farm); much faster than primary.
- Pioneer species
- The first organisms to colonize a disturbed area (lichens, mosses, fast-growing weeds), which begin building soil.
- Climax community
- A relatively stable, mature community in equilibrium with the local climate at the end of succession.
- Resilience (ecological)
- The ability of an ecosystem to recover after a disturbance; higher biodiversity generally increases resilience.
- Indicator species
- A species whose health reflects the condition of its environment (e.g., amphibians for water quality).
- Specialist species
- A species with a narrow niche and specific requirements (e.g., the panda); vulnerable to environmental change.
- Generalist species
- A species with a broad niche that can use many resources (e.g., raccoons); adaptable to change.
- Ecological niche
- The role a species plays in its ecosystem — what it eats, where it lives, and how it interacts with others.
- Island biogeography
- The theory that larger islands closer to the mainland support more species than small, remote ones.
- Mass extinction
- A period when a large fraction of Earth's species go extinct over a short geologic time; Earth has had five.
- Mutualism
- A symbiotic relationship in which both species benefit (e.g., bees and flowers).
- Commensalism
- A relationship in which one species benefits and the other is unaffected (e.g., barnacles on a whale).
- Parasitism
- A relationship in which one species (the parasite) benefits at the host's expense.
- Predator-prey relationship
- An interaction where one organism hunts and eats another; their populations rise and fall in linked cycles.
- Ecological tolerance
- The range of conditions (temperature, pH, salinity) an organism can survive; outside it, the organism cannot live.
- Carrying capacity (K)
- The maximum population size an environment can support indefinitely given its resources.
- Exponential growth
- J-shaped growth that occurs with unlimited resources; the population grows by a constant percent each interval.
- Logistic growth
- S-shaped growth that slows as a population nears its carrying capacity and limiting factors increase.
- Density-dependent factor
- A limiting factor that intensifies as population density rises: food shortage, disease, predation, competition.
- Density-independent factor
- A limiting factor that affects a population regardless of its density: weather, natural disasters, climate.
- r-selected species
- Species with many small offspring, little parental care, and short lives (insects, weeds); boom-and-bust populations.
- K-selected species
- Species with few large offspring, high parental care, and long lives (elephants, oaks); stable near carrying capacity.
- Survivorship curve Type I
- Most individuals survive to old age then die quickly (humans, large mammals).
- Survivorship curve Type III
- Most individuals die young in huge numbers; few survive to adulthood (fish, insects, plants).
- Population growth rate formula
- Growth rate = (births − deaths + immigration − emigration). For a simple rate: (births − deaths) ÷ 10 per 1,000.
- Rule of 70
- Doubling time (years) ≈ 70 ÷ the percent growth rate. A 2% growth rate doubles in about 35 years.
- Demographic transition model
- A four-stage model of how birth and death rates fall as a country develops.
- Total fertility rate (TFR)
- The average number of children a woman has in her lifetime; about 2.1 is replacement level in developed nations.
- Replacement-level fertility
- The TFR (~2.1) at which a population exactly replaces itself, leading to stable size over time.
- Age-structure diagram
- A graph of a population by age and sex; a wide base predicts rapid growth, a narrow base predicts decline.
- Infant mortality rate
- The number of infant deaths per 1,000 live births; a key indicator of a country's development.
- Doubling time
- The time it takes a population to double in size, estimated with the rule of 70.
- Limiting factor
- Anything that restricts the size of a population, such as food, water, space, or predators.
- Population crash (die-off)
- A sharp drop in numbers after a population overshoots its carrying capacity.
- Biotic potential
- The maximum reproductive rate of a population under ideal conditions.
- Immigration vs. emigration
- Immigration is individuals moving INTO a population; emigration is individuals moving OUT. Both change population size.
- Demographic momentum
- Continued population growth after fertility falls to replacement level, because of a large young population.
- Plate tectonics
- The movement of Earth's crustal plates, which causes earthquakes, volcanoes, and mountain-building at boundaries.
- Soil horizon
- A layer in a soil profile, from O (organic litter) and A (topsoil) at the top down to C and bedrock.
- Soil texture
- The proportion of sand, silt, and clay in soil; it controls water and nutrient holding. Loam is best for farming.
- Loam
- A balanced mix of sand, silt, and clay that holds water and nutrients well — ideal agricultural soil.
- Troposphere
- The lowest atmospheric layer where weather occurs and most air mass and water vapor are found.
- Stratosphere
- The atmospheric layer above the troposphere that contains the protective ozone layer.
- Hadley cell
- A global circulation cell where air rises at the equator (wet) and sinks around 30° latitude (dry, where deserts form).
- Coriolis effect
- The deflection of moving air and water by Earth's rotation, which shapes global wind belts and ocean currents.
- El Niño
- A periodic warming of the eastern Pacific that weakens trade winds, suppresses upwelling, and shifts global weather.
- La Niña
- The cool phase of ENSO with stronger trade winds and cooler eastern Pacific water — the opposite of El Niño.
- Upwelling
- The rise of cold, nutrient-rich water from the deep ocean to the surface, supporting productive fisheries.
- Aquifer
- An underground layer of permeable rock or sediment that stores and transmits groundwater.
- Watershed
- The land area that drains into a particular river, lake, or other body of water.
- Albedo
- The fraction of sunlight a surface reflects; ice and snow are high (reflective), dark land and ocean are low (absorbent).
- Milankovitch cycles
- Long-term natural variations in Earth's orbit and tilt that drive ice ages — a natural, not human, climate driver.
- Weathering
- The physical and chemical breakdown of rock at Earth's surface, the first step in soil formation.
- Erosion
- The transport of weathered soil and rock by water, wind, or ice; worsened by removing vegetation.
- Salinization
- The buildup of salts in soil from irrigation in dry climates, which reduces crop productivity.
- Permeability
- How easily water and air move through soil or rock; sandy soil is highly permeable, clay is not.
- Thermohaline circulation
- The global ocean conveyor belt driven by differences in water temperature and salinity (density).
- Rain shadow
- The dry region on the leeward side of a mountain, where descending air has lost its moisture.
- Earthquake / Richter scale
- Ground shaking from plate movement; the Richter (or moment magnitude) scale measures its energy.
- Tragedy of the commons
- The overuse of a shared, unregulated resource because individuals benefit while the cost of depletion is shared by all.
- Green Revolution
- The 20th-century increase in crop yields through fertilizer, irrigation, pesticides, and high-yield seed varieties.
- Contour plowing
- Plowing across a slope along its contour lines to reduce soil erosion and water runoff.
- No-till farming
- Planting without plowing, which keeps soil and moisture in place and stores carbon.
- Crop rotation
- Alternating the crops grown in a field to restore soil nitrogen and break pest cycles.
- Integrated pest management (IPM)
- Combining biological, mechanical, and limited chemical controls to manage pests with less pesticide.
- Monoculture
- Growing a single crop over a large area; efficient but vulnerable to pests and hard on soil.
- Overgrazing
- Letting livestock eat vegetation faster than it regrows, leading to soil erosion and desertification.
- Desertification
- The degradation of once-productive land into desert, often from overgrazing, deforestation, or drought.
- CAFO
- Concentrated Animal Feeding Operation; raises many animals in a small area and generates large amounts of waste.
- Sustainable yield
- Harvesting a renewable resource no faster than it can replenish itself.
- Surface (strip) mining
- Removing surface layers to reach a shallow deposit; cheaper but highly destructive to the land.
- Mountaintop removal
- A form of surface mining that blasts off mountain tops to reach coal, burying streams with debris.
- Subsurface mining
- Tunneling underground to reach deep deposits; costlier and more dangerous than surface mining.
- Mine reclamation
- Restoring land after mining by regrading, replacing topsoil, and replanting vegetation.
- Overfishing
- Catching fish faster than populations can reproduce, leading to collapse; managed with catch limits.
- Aquaculture
- Farming fish and shellfish; boosts food supply but can pollute water and spread disease.
- Clear-cutting
- Removing all trees in an area at once; cheap but causes erosion and habitat loss.
- Urban sprawl
- The spread of low-density development outward from a city, consuming farmland and increasing car dependence.
- Ecological footprint
- The amount of productive land and water needed to supply a person's resources and absorb their waste.
- Tragedy fix: regulation/ownership
- Solutions to the tragedy of the commons: government quotas/regulation or assigning private ownership.
- Buffer strip
- A vegetated zone along a waterway that filters runoff and reduces fertilizer and sediment pollution.
- Nonrenewable energy
- Energy from finite stocks formed over millions of years: coal, oil, natural gas, and nuclear fuel.
- Renewable energy
- Energy from sources that replenish on a human timescale: solar, wind, hydro, geothermal, and biomass.
- Fossil fuels
- Coal, oil, and natural gas formed from buried ancient organisms; burning them releases CO₂ and pollutants.
- Coal
- The dirtiest fossil fuel; abundant and cheap but releases the most CO₂, SO₂, mercury, and particulates.
- Natural gas
- The cleanest-burning fossil fuel; releases less CO₂ than coal but leaks methane, a potent greenhouse gas.
- Nuclear fission
- Splitting uranium-235 atoms to release heat that boils water to spin a turbine; no CO₂ but radioactive waste.
- Control rods
- Rods in a nuclear reactor that absorb neutrons to slow or stop the fission chain reaction.
- Nuclear meltdown
- Overheating that melts a reactor core and can release radiation (e.g., Chernobyl, Fukushima).
- Solar (photovoltaic) energy
- Converting sunlight directly into electricity with solar cells; clean but intermittent.
- Wind energy
- Using turbines to convert wind into electricity; clean and renewable but intermittent and location-dependent.
- Hydroelectric power
- Generating electricity from flowing water, usually behind a dam; reliable but disrupts rivers and fish.
- Geothermal energy
- Using Earth's internal heat to generate electricity or heat buildings; steady and clean but location-limited.
- Biomass energy
- Burning organic material (wood, crops, waste) for energy; renewable but releases CO₂ and air pollutants.
- Energy conservation
- Reducing energy use through efficiency and behavior; the cheapest and cleanest 'source' of energy.
- Energy efficiency
- Getting more useful output from each unit of energy input (e.g., LED bulbs, hybrid cars).
- Cogeneration (CHP)
- Capturing waste heat from electricity generation to use for heating, raising overall efficiency.
- Fuel cell
- A device that produces electricity from hydrogen and oxygen, emitting only water; clean but costly.
- Cap and trade
- A market policy that sets a pollution limit and lets companies buy and sell emission allowances.
- Net energy yield
- The usable energy from a source after subtracting the energy needed to produce it.
- Tar sands / oil shale
- Unconventional fossil fuels that are energy- and water-intensive to extract and process.
- Fracking (hydraulic fracturing)
- Injecting high-pressure fluid to crack rock and release natural gas or oil; can contaminate groundwater.
- Three Gorges / dam trade-offs
- Large dams provide renewable power and flood control but flood habitat, displace people, and block fish.
- Primary air pollutant
- A pollutant emitted directly into the air (CO, SO₂, NOₓ, particulates, VOCs).
- Secondary air pollutant
- A pollutant that forms in the atmosphere from reactions of primary pollutants (e.g., ground-level ozone).
- Photochemical smog
- A brownish haze of ground-level ozone formed when sunlight reacts with NOₓ and VOCs on hot, sunny days.
- Ground-level ozone
- A harmful secondary pollutant in smog that damages lungs and crops — 'bad' ozone, nearby.
- Thermal inversion
- A layer of warm air over cooler air near the ground that traps pollutants and worsens smog.
- Particulate matter (PM)
- Tiny solid or liquid particles in the air (soot, dust); PM2.5 lodges deep in the lungs and harms health.
- Acid rain
- Precipitation acidified by sulfuric and nitric acids formed when SO₂ and NOₓ react with water in the air.
- Sulfur dioxide (SO₂)
- A pollutant from burning coal that contributes to acid rain and respiratory problems.
- Nitrogen oxides (NOₓ)
- Pollutants from combustion that form smog and acid rain and contribute to ground-level ozone.
- Carbon monoxide (CO)
- A colorless, odorless gas from incomplete combustion that binds hemoglobin and reduces oxygen delivery.
- Volatile organic compounds (VOCs)
- Carbon compounds that evaporate easily (from fuels, paints, solvents) and help form photochemical smog.
- Clean Air Act
- The U.S. law that regulates air pollutants and sets National Ambient Air Quality Standards.
- Scrubber
- A smokestack device that removes SO₂ and particulates from exhaust gases before they reach the air.
- Catalytic converter
- A device on cars that converts CO, NOₓ, and VOCs in exhaust into less harmful gases.
- Electrostatic precipitator
- A device that uses an electric charge to remove particulates from smokestack exhaust.
- Radon
- A naturally occurring radioactive gas that seeps into buildings from soil; a leading cause of indoor lung cancer.
- Asbestos
- A fibrous mineral once used as insulation; inhaling its fibers causes lung disease and cancer.
- Sick building syndrome
- Health problems linked to poor indoor air quality in a building, from VOCs, mold, and poor ventilation.
- Industrial (gray) smog
- Smog from burning coal and oil, high in sulfur dioxide and particulates; common in industrial cities.
- Indoor air pollution
- Pollutants inside buildings (radon, CO, VOCs, mold, asbestos), often a greater health risk than outdoor air.
- Noise pollution
- Unwanted or harmful sound that can cause hearing loss and stress in humans and disrupt wildlife.
- Lead (Pb) as a pollutant
- A toxic metal once in gasoline and paint that harms the nervous system, especially in children.
- Eutrophication
- Nutrient enrichment of water (nitrogen, phosphorus) that triggers algal blooms; decay then depletes oxygen.
- Cultural eutrophication
- Human-accelerated eutrophication from fertilizer runoff, sewage, and detergents.
- Dead zone (hypoxia)
- An area of water with too little dissolved oxygen to support most life, caused by eutrophication.
- Point-source pollution
- Pollution from a single identifiable source, such as a factory discharge pipe — easier to regulate.
- Nonpoint-source pollution
- Diffuse pollution from many sources, such as farm and street runoff — harder to control.
- Biochemical oxygen demand (BOD)
- The amount of oxygen decomposers use to break down organic waste in water; high BOD starves water of oxygen.
- Bioaccumulation
- The buildup of a persistent toxin in one organism's tissues over its lifetime.
- Biomagnification
- The increase in a toxin's concentration at each higher trophic level, so top predators carry the most.
- DDT
- A persistent pesticide that biomagnified up food chains and thinned the eggshells of birds of prey like eagles.
- Mercury (as a pollutant)
- A toxic metal from coal burning and mining that biomagnifies in fish and harms the nervous system.
- Persistent organic pollutant (POP)
- A long-lasting, fat-soluble toxic chemical (DDT, PCBs) that biomagnifies and resists breakdown.
- Dose-response curve
- A graph showing how the harmful effect of a substance increases with the dose received.
- LD50
- The dose of a toxin that is lethal to 50% of a test population; a standard measure of toxicity.
- Sanitary landfill
- A lined and covered site for solid waste, designed to protect groundwater from leachate.
- Leachate
- Liquid that drains through a landfill, picking up contaminants that can pollute groundwater.
- Incineration
- Burning solid waste to reduce its volume and generate energy; releases air pollutants and ash.
- The 3 Rs (reduce, reuse, recycle)
- The waste hierarchy: reduce consumption first, then reuse, then recycle before disposal.
- Sewage treatment (primary)
- The first stage that physically removes solids from wastewater by screening and settling.
- Sewage treatment (secondary)
- The biological stage that uses bacteria to break down dissolved organic waste in wastewater.
- Thermal pollution
- The release of heated water (often from power plants) that lowers dissolved oxygen and harms aquatic life.
- Oil spill effects
- Oil coats organisms and feathers, smothers habitats, and is hard to clean; harms birds, mammals, and fisheries.
- Clean Water Act
- The U.S. law that regulates pollutant discharges into surface waters and sets water-quality standards.
- Greenhouse effect
- Natural warming when greenhouse gases (CO₂, methane, water vapor, nitrous oxide) absorb and re-emit Earth's infrared heat.
- Enhanced greenhouse effect
- The extra warming caused by human emissions of greenhouse gases above natural levels.
- Carbon dioxide (CO₂)
- The most abundant human-released greenhouse gas, mainly from burning fossil fuels and deforestation.
- Methane (CH₄)
- A greenhouse gas that traps far more heat per molecule than CO₂ but is less abundant; from livestock, landfills, and gas leaks.
- Global warming
- The long-term rise in Earth's average surface temperature driven by the enhanced greenhouse effect.
- Climate change
- Long-term shifts in temperature and weather patterns, largely driven by human greenhouse-gas emissions.
- Ocean acidification
- The drop in ocean pH as CO₂ dissolves to form carbonic acid, harming coral and shell-building organisms.
- Sea-level rise
- The rise in ocean levels from melting land ice and the thermal expansion of warming seawater.
- Positive feedback loop
- A change that amplifies itself; melting ice lowers albedo, absorbing more heat and melting more ice.
- Albedo feedback
- As reflective ice melts it exposes dark land/ocean, which absorbs more heat and accelerates warming.
- Ozone depletion
- Thinning of the stratospheric ozone layer by chlorine from CFCs, letting more harmful UV reach the surface.
- Chlorofluorocarbons (CFCs)
- Chemicals once used in refrigerants and aerosols that release ozone-destroying chlorine in the stratosphere.
- Montreal Protocol
- The 1987 treaty that phased out CFCs and is allowing the ozone layer to slowly recover — a major success.
- Stratospheric vs. ground-level ozone
- Stratospheric ozone protects life from UV ('good up high'); ground-level ozone is a pollutant ('bad nearby').
- Invasive species
- A non-native species that spreads rapidly and harms an ecosystem, often lacking natural predators (zebra mussels, kudzu).
- HIPPCO
- The drivers of biodiversity loss: Habitat destruction, Invasive species, Pollution, Population, Climate change, Overexploitation.
- Habitat fragmentation
- Breaking a large habitat into smaller, isolated patches, which reduces biodiversity and species movement.
- Endangered species
- A species at serious risk of extinction throughout all or much of its range.
- Endangered Species Act
- The U.S. law that protects threatened and endangered species and their critical habitat.
- Tipping point
- A threshold beyond which a system shifts abruptly and often irreversibly to a new state (e.g., a dying coral reef).
- Kyoto Protocol / Paris Agreement
- International agreements to reduce greenhouse-gas emissions and limit global warming.
- Carbon footprint
- The total greenhouse gases produced by a person, product, or activity, expressed as CO₂ equivalent.
- Carbon sequestration
- Capturing and storing atmospheric CO₂ in forests, soils, oceans, or underground to slow warming.
- Deforestation and climate
- Clearing forests removes a carbon sink and releases stored carbon, worsening global warming.